Besania

Besania Temporal range: Anisian–Ladinian[1] PreꞒ Ꞓ O S D C P T J K Pg N
Fossil of Besania sp. at Civic Museum of Fossils of Besano
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Subclass:	Neopterygii
Genus:	†Besania Brough, 1939
Type species
†Besania micrognathus Brough, 1939
Other species
†B. schaufelbergerae Herzog & Bürgin, 2005

Besania is an extinct genus of prehistoric marine ray-finned fish^[1] that lived during the Anisian and Ladinian ages of the Middle Triassic epoch in what is now southern/southeastern Switzerland and northern Italy.^[2] Fossils were recovered from the Besano Formation of Monte San Giorgio area (Swiss-Italian borderland)^[3] and the Prosanto Formation of canton Graubünden, Switzerland.^[4]

Initially considered a perleidiform or luganoiiform, it is now thought to be an indeterminate "halecostome" or basal neopterygian.^[4] There are two known species, Besania micrognathus (type species) and B. schaufelbergerae. Both are quite small (31 mm (1.2 in)–44 mm (1.7 in) in standard length).

The genus is named after the Italian town of Besano, Lombardy, near Monte San Giorgio.

• Prehistoric fish
• List of prehistoric bony fish

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Neopterygii

Kingdom: Animalia Phylum: Chordata Subphylum: Vertebrata Class: Actinopterygii Clade: Actinopteri
Actinopterygii see Actinopterygii Neopterygii see below↓ Neopterygii

Neopterygii	†Besania †Boreichthys †Caelatichthys †Cephaloxenus †Chrotichthys †Dollopterus †Frodoichthys †Gimlichthys †Habroichthys †Hulettia †Pristisomus †Tungusichthys †Venusichthys †Zeuchthiscus †Bobasatraniiformes? †Guildayichthyidae? †Pholidopleuriformes? Arctosomus Australosomus Gracilignathichthys Macroaethes Pholidopleurus †Redfieldiiformes? Atopocephala Brookvalia Calaichthys Cionichthys Daedalichthys Denwoodichthys Dictyopyge Endemichthys Geitonichthys Helichthys Hiascoactinus Ischnolepis Lasalichthys Leighiscus Mauritanichthys Molybdichthys Pacorichthys Phlyctaenichthys Redfieldius Sakamenichthys Schizurichthys Sinkiangichthys? †Platysiagidae Caelatichthys? Helmolepis Platysiagum †Polzbergiiformes Hydropessum? Cleithrolepididae Cleithrolepidina Cleithrolepis Polzbergiidae Felberia Luopingichthys? Polzbergia Serrolepis Stoppania †Colobodontidae Chaohuperleidus? Colobodus Crenilepis Dactylolepis? Engycolobodus? Feroxichthys Gabanellia? Nephrotus? †Perleididae Aetheodontus? Alvinia? Chaohuperleidus? Ctenognathichthys? Daninia Diandongperleidus? Dipteronotus? Endennia? Eoperleidus Fuyuanperleidus? Luopingperleidus? Manlietta Megaperleidus? Meidiichthys Meridensia? Moradebrichthys? Paraperleidus Peltoperleidus? Perleidus Plesiofuro? Plesioperleidus Procheirichthys? Teffichthys? Tripelta †Louwoichthyiformes Peltoperleidus? Louwoichthyidae Ctenognathichthys? Louwoichthys Luopingichthys? Pseudobeaconiidae Anatoia Caminchaia Dipteronotus? Echentaia Mendocinichthys Pasambaya Pseudobeaconia †Luganoiiformes Altisolepis? Fuyuanperleidus? Luganoia †Peltopleuriformes Habroichthys? Peltopleuridae Altisolepis? Nannolepis Peltopleurus Placopleurus Thoracopteridae Gigantopterus Italopterus Peripeltopleurus Potanichthys Thoracopterus Wushaichthys †Dapediidae Aetheolepis Dandya Dapedium Hemicalypterus Heterostrophus Paradapedium Sargodon Scopulipiscis Tetragonolepis †Pycnodontiformes see Pycnodontiformes Holostei Ginglymodi see Ginglymodi includes gars Halecomorphi see Halecomorphi includes bowfin Teleostei see Teleostei includes the majority of modern fish

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