Microphysidae

Family of true bugs

Microphysidae Temporal range: Late Cretaceous–Present PreꞒ Ꞓ O S D C P T J K Pg N

male Loricula sp. (<2 mm) from Cologne, Germany
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hemiptera
Suborder:	Heteroptera
Infraorder:	Cimicomorpha
Superfamily:	Miroidea
Family:	Microphysidae Dohrn, 1859
Genera
See text

The Microphysidae are a very small family of true bugs, comprising only 5 extant genera.

Systematics

female *Loricula* sp. (<2 mm) from Cologne, Germany

Until recently, many authors considered these bugs to belong within the family Anthocoridae.^{[citation needed]} A 2022 molecular phylogenetics study groups them with Joppeicidae in the proposed clade Microphysoidea.^[1] A 2023 study recovers them instead as relatively basal Cimiciformes.^[2]

The following genera belong to this family:

Subfamily Microphysinae

Genus Chinaola Blatchley
Genus Loricula Curtis (= Microphysa)
Genus Mallochiola Bergroth
Genus Myrmedobia Bärensprung
Genus †Myrmericula Popov
Genus †Popovophysa McKellar & Engel 2011, Canadian amber, Campanian
Genus †Tytthophysa Popov & Herczek 2009 Baltic amber, Eocene

Subfamily Ciorullinae

Genus Ciorulla Péricart

References

^ Ye, Fei; Kment, Petr; Rédei, Dávid; Luo, Jiu-Yang; Wang, Yan-Hui; Kuechler, Stefan M.; Zhang, Wei-Wei; Chen, Ping-Ping; Wu, Hao-Yang; Wu, Yan-Zhuo; Sun, Xiao-Ya; Ding, Lu; Wang, Yue-Ran; Xie, Qiang (2022). "Diversification of the phytophagous lineages of true bugs (Insecta: Hemiptera: Heteroptera) shortly after that of the flowering plants". Cladistics. 38 (4): 403–428. doi:10.1111/cla.12501. PMID 35349192.
^ Jung, Sunghoon; Kim, Junggon; Balvín, Ondřej; Yamada, Kazutaka (2023). "Molecular Phylogeny of Cimicoidea (Heteroptera: Cimicomorpha) Revisited: Increased Taxon Sampling Reveals Evolution of Traumatic Insemination and Paragenitalia". Insects. 14 (3): 267. doi:10.3390/insects14030267. PMC 10051671. PMID 36975952.

Extant Hemiptera families

Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Subclass: Pterygota
Infraclass: Neoptera
Superorder: Paraneoptera

Suborder Auchenorrhyncha

Cicadomorpha

Cercopoidea (froghoppers)	Aphrophoridae Cercopidae Clastopteridae Epipygidae Machaerotidae
Cicadoidea (cicadas)	Cicadidae Tettigarctidae (hairy cicadas)
Membracoidea	Aetalionidae Cicadellidae (leafhoppers) Melizoderidae Membracidae (typical treehoppers, thorn bugs) Myerslopiidae

Fulgoromorpha
(planthoppers)

Fulgoroidea	Acanaloniidae Achilidae Achilixiidae Cixiidae Delphacidae Derbidae Dictyopharidae Eurybrachidae Flatidae Fulgoridae (lanternflies) Gengidae Hypochthonellidae Issidae Kinnaridae Lophopidae Meenoplidae Nogodinidae Ricaniidae Tettigometridae Tropiduchidae

Suborder Sternorrhyncha

Aleyrodoidea	Aleyrodidae (whiteflies)
Aphidoidea (aphids)	Aphididae
Coccoidea (scale insects)	Aclerdidae Asterolecaniidae (pit scales) Beesoniidae (beesoniids) Carayonemidae (carayonemids) Cerococcidae (ornate pit scales) Coccidae (soft scales) Conchaspididae Dactylopiidae (cochineals) Diaspididae (armored scales) Eriococcidae (felt scales) Halimococcidae (pupillarial palm scales) Kermesidae Kerriidae (lac scales) Lecanodiaspididae (false pit scales) Margarodidae (cottony cushion scales, giant coccids, ground pearls) Micrococcidae (Mediterranean scales) Monophlebidae (giant scales) Ortheziidae (ensign scales) Phenacoleachiidae (phenacoleachiids) Phoenicococcidae (palm scales) Pseudococcidae (mealybugs) Putoidae (giant mealybugs) Stictococcidae (stictococcids)
Phylloxeroidea	Adelgidae (woolly conifer aphids) Phylloxeridae (phylloxerans)
Psylloidea	Aphalaridae Calophyidae Carsidaridae Homotomidae Phacopteronidae Psyllidae (jumping plant lice) Triozidae

Suborder Heteroptera (with Coleorrhyncha)

Dipsocoromorpha	Ceratocombidae Dipsocoridae Hypsipterygidae Schizopteridae Stemmocryptidae

Enicocephalomorpha

Enicocephaloidea	Aenictopecheidae Enicocephalidae (unique-headed bugs, gnat bugs)

Gerromorpha
(semiaquatic bugs)

Gerroidea	Gerridae (water striders) Hermatobatidae Veliidae (riffle bugs)
Hebroidea	Hebridae (velvet water bugs)
Hydrometroidea	Hydrometridae (marsh treaders or water measurers) Macroveliidae Paraphrynoveliidae
Mesovelioidea (water treaders)	Madeoveliidae Mesoveliidae

Leptopodomorpha	Saldidae (shore bugs) Leptopodidae (spiny shore bugs) Omaniidae Aepophilidae

Nepomorpha
(true water bugs)

Corixoidea	Corixidae (water boatmen)
Nepoidea	Belostomatidae (giant water bugs) Nepidae (water scorpions, needle bugs)
Ochteroidea	Gelastocoridae (toad bugs) Ochteridae (velvety shore bugs)
Aphelocheiroidea	Aphelocheiridae Potamocoridae
Naucoroidea	Naucoridae (creeping water bugs)
Notonectoidea	Notonectidae (backswimmers)
Pleoidea	Helotrephidae Pleidae (pygmy backswimmers)

Peloridiomorpha (Coleorrhyncha)	Peloridiidae

Cimicomorpha

Cimicoidea

Anthocoridae (minute pirate bugs or flower bugs)
Cimicidae (bed bugs, bat bugs)
Curaliidae (Curalium cronini)
Joppeicidae
Lasiochilidae
Lyctocoridae
Medocostidae
Microphysidae
Miridae (plant bugs, leaf bugs, grass bugs)
Nabidae (damsel bugs)
Pachynomidae
Plokiophilidae
Polyctenidae (old world bat bugs)
Reduviidae (assassin bugs, wheel bugs, thread-legged bugs)
Thaumastocoridae (royal palm bugs)
Tingidae (lace bugs)
Velocipedidae

Pentatomomorpha

Aradoidea	Aradidae (flat bugs) Termitaphididae (termite bugs)
Pentatomoidea (shield bugs)	Urostylididae Acanthosomatidae (shield bugs) Tessaratomidae (giant shield bugs and relatives) Dinidoridae Cydnidae (burrowing bugs) Thaumastellidae Parastrachiidae Thyreocoridae (includes ebony bugs) Lestoniidae Phloeidae Scutelleridae (jewel bugs or metallic shield bugs) Plataspidae Pentatomidae (stink bugs) Canopidae Megarididae
Coreoidea	Alydidae (broad-headed bugs) Coreidae (squash bugs, leaf-footed bugs) Hyocephalidae Rhopalidae (scentless plant bugs) Stenocephalidae
Lygaeoidea	Artheneidae Berytidae (stilt bugs) Blissidae Colobathristidae Cryptorhamphidae Cymidae Geocoridae Henicocoridae Heterogastridae Idiostolidae Lygaeidae (milkweed bugs, true seed bugs) Malcidae Ninidae Oxycarenidae Pachygronthidae Rhyparochromidae (atypical seed bugs) Piesmatidae (ash-grey leaf bugs)
Pyrrhocoroidea	Pyrrhocoridae (red bugs, cotton stainers) Largidae (bordered plant bugs)

Note: Coleorrhyncha are a different clade from Heteroptera. Heteroptera with Coleorrhyncha were referred to as Prosorrhyncha.

Taxon identifiers
Microphysidae	Wikidata: Q1427608 Wikispecies: Microphysidae BioLib: 17139 BOLD: 365828 BugGuide: 333590 CoL: CRS EoL: 610 EPPO: 1MPHYF Fauna Europaea: 12829 Fauna Europaea (new): 9f5e4735-c22f-4dcd-bd81-83f0c4cf32bc GBIF: 7890 iNaturalist: 174329 IRMNG: 101487 ITIS: 104409 NBN: NBNSYS0000160423 NCBI: 236447 Open Tree of Life: 907378 Paleobiology Database: 217451